Penny et al. 2001

Penny, David and McComish, Bennett J. and Charleston, Michael A. and Hendy, Michael D. 2001. Mathematical Elegance with Biochemical Realism: The Covarion Model of Molecular Evolution. Journal of Molecular Evolution 53. 711–723.

@article{jacquesestimative-penny01covarion,
  author  = {Penny, David and McComish, Bennett J. and Charleston, Michael A. and Hendy, Michael D.},
  journal = {Journal of Molecular Evolution},
  number  = {6},
  pages   = {711–723},
  title   = {Mathematical Elegance with Biochemical Realism: The Covarion Model of Molecular Evolution},
  volume  = {53},
  year    = {2001}
}